Python implementation of the DarkDiv R package for calculating dark diversity from co-occurrence data
Project description
PyDarkDiv
Python implementation of the DarkDiv R package for calculating dark diversity from species co-occurrence data.
Overview
PyDarkDiv estimates dark diversity - the unobserved portion of a site's species pool consisting of species that could potentially occur but are currently absent. This package implements the methods described in Carmona & Pärtel (2021).
Based on: DarkDiv R package by Carlos P. Carmona and Meelis Pärtel
Installation
pip install pydarkdiv
Quick Start
import pandas as pd
import pydarkdiv as pdd
# Load your species data (sites × species matrix)
data = pd.read_csv('species_data.csv', index_col=0)
# Calculate dark diversity
dd = pdd.DarkDiv(data, method='Hypergeometric')
# Get results as DataFrames
dfs = dd.to_dataframes()
dark_diversity = dfs['dark']
species_pool = dfs['pool']
# Summary statistics
dark_richness = (dark_diversity > 0.5).sum(axis=1)
print(f"Average dark diversity: {dark_richness.mean():.1f} species")
Methods
- Hypergeometric (recommended): Compares observed vs. expected co-occurrences
- RawBeals: Beals smoothing for occurrence probabilities
- Favorability: Favorability-corrected Beals values
- ThresholdBeals: Binary predictions using thresholds
Usage Examples
See usage_examples.py for detailed examples including:
- Basic usage
- Method comparisons
- Reference data usage
- Abundance weighting
Requirements
- Python ≥ 3.8
- numpy ≥ 1.21.0
- pandas ≥ 1.3.0
- scipy ≥ 1.7.0
Reference
Carmona, C.P. & Pärtel, M. (2021). Estimating probabilistic site-specific species pools and dark diversity from co-occurrence data. Global Ecology and Biogeography, 30(1), 316-326. DOI link
License
MIT License
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